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| Holder | Type | Held (Actual) | % of Outstanding | Market Value | Date Reported |
|---|---|---|---|---|---|
| RA Capital Management, L.P. | Institution | 6,116,140 | 12.59% | US$ 2.23M | 2026-05-12 |
| RA Capital Healthcare Fund, L.P. | Institution | 5,709,858 | 11.75% | US$ 2.08M | 2026-05-12 |
| Longwood Fund III, L.P. | Institution | 1,677,082 | 3.45% | US$ 612.13k | 2025-12-31 |
| Longwood Fund | Institution | 1,677,082 | 3.45% | US$ 612.13k | 2025-12-31 |
| Vanguard Capital Management, LLC | Institution | 1,217,976 | 2.51% | US$ 444.56k | 2026-03-31 |
| Lynx1 Capital Management LP | Institution | 657,786 | 1.35% | US$ 240.09k | 2026-03-31 |
| UBS Oncology Impact Fund | Institution | 655,471 | 1.35% | US$ 239.25k | 2026-05-12 |
| BioImpact Capital LLC | Institution | 655,471 | 1.35% | US$ 239.25k | 2026-05-12 |
| Dr. Daniel Hicklin | Insider | 647,047 | 1.33% | US$ 236.17k | 2025-04-14 |
| Sphera Funds Management Ltd | Institution | 626,810 | 1.29% | US$ 228.79k | 2026-03-31 |
| DC Investments Management, LLC | Institution | 589,102 | 1.21% | US$ 215.02k | 2026-03-31 |
| Squadron Capital Management Llc | Institution | 563,535 | 1.16% | US$ 205.69k | 2026-03-31 |
| Renaissance Technologies LLC | Institution | 516,218 | 1.06% | US$ 188.42k | 2026-03-31 |
| MPM BioImpact LLC | Institution | 482,776 | 0.99% | US$ 176.21k | 2026-06-03 |
| RA Capital Nexus Fund II, L.P. | Institution | 406,282 | 0.84% | US$ 148.29k | 2026-05-12 |
| Geode Capital Management, LLC | Institution | 389,487 | 0.80% | US$ 142.16k | 2026-03-31 |
| MPM Bioventures 2014, L.P. | Institution | 348,364 | 0.72% | US$ 127.15k | 2026-06-03 |
| Merrill Lynch, Pierce, Fenner & Smith Incorporated | Institution | 241,034 | 0.50% | US$ 87.98k | 2026-03-31 |
| Vanguard Fiduciary Trust Company | Institution | 157,726 | 0.32% | US$ 57.57k | 2026-03-31 |
| Millennium Management LLC | Institution | 132,050 | 0.27% | US$ 48.20k | 2026-03-31 |
| State Street Global Advisors, Inc. | Institution | 119,168 | 0.25% | US$ 43.50k | 2026-03-31 |
| BlackRock, Inc. | Institution | 101,608 | 0.21% | US$ 37.09k | 2026-03-31 |
| Northern Trust Global Investments | Institution | 86,851 | 0.18% | US$ 31.70k | 2026-03-31 |
| Alyeska Investment Group, L.P. | Institution | 86,554 | 0.18% | US$ 31.59k | 2026-03-31 |
| Jane Street Group, LLC | Institution | 82,836 | 0.17% | US$ 30.24k | 2026-03-31 |
| Bridgeway Capital Management, LLC | Institution | 77,000 | 0.16% | US$ 28.11k | 2026-03-31 |
| MPM Oncology Innovations Fund LP | Institution | 71,451 | 0.15% | US$ 26.08k | 2026-06-03 |
| BBVA Asset Management, S.A., S.G.I.I.C | Institution | 70,032 | 0.14% | US$ 25.56k | 2025-06-30 |
| UBS Asset Management AG | Institution | 66,857 | 0.14% | US$ 24.40k | 2026-03-31 |
| XTX Markets Limited | Institution | 50,929 | 0.10% | US$ 18.59k | 2026-03-31 |
| The Goldman Sachs Group, Inc. | Institution | 38,266 | 0.08% | US$ 13.97k | 2026-03-31 |
| Morgan Stanley | Institution | 33,799 | 0.07% | US$ 12.34k | 2026-03-31 |
| Simplex Trading, LLC | Institution | 32,526 | 0.07% | US$ 11.87k | 2026-03-31 |
| Verdad Advisers, L.P. | Institution | 29,760 | 0.06% | US$ 10.86k | 2026-03-31 |
| Bank of America Corporation | Institution | 23,549 | 0.05% | US$ 8.60k | 2026-03-31 |
| Raymond James Financial, Inc. | Institution | 19,778 | 0.04% | US$ 7.22k | 2026-03-31 |
| Hudson River Trading LLC | Institution | 17,861 | 0.04% | US$ 6.52k | 2026-03-31 |
| Dr. Briggs Duncan W. Morrison | Insider | 15,523 | 0.03% | US$ 5.67k | 2025-04-14 |
| Certuity, LLC | Institution | 15,000 | 0.03% | US$ 5.48k | 2026-03-31 |
| Delta Investment Management, LLC | Institution | 14,925 | 0.03% | US$ 5.45k | 2026-03-31 |
| Dr. Michael Atkins | Insider | 14,419 | 0.03% | US$ 5.26k | 2025-04-14 |
| Barclays PLC | Institution | 13,235 | 0.03% | US$ 4.83k | 2026-03-31 |
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