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| Holder | Type | Held (Actual) | % of Outstanding | Market Value | Date Reported |
|---|---|---|---|---|---|
| Flagship Pioneering, Inc | Institution | 12,674,120 | 21.59% | US$ 51.71M | 2026-03-31 |
| Flagship Ventures Fund V, L.P. | Institution | 9,330,878 | 15.89% | US$ 38.07M | 2026-01-13 |
| FMR LLC | Institution | 6,047,198 | 10.30% | US$ 24.67M | 2026-03-31 |
| BVF Partners L.P. | Institution | 5,842,000 | 9.95% | US$ 23.84M | 2026-03-31 |
| Deerfield Partners, LP | Institution | 5,139,523 | 8.75% | US$ 20.97M | 2026-03-31 |
| Deerfield Management Company, L.P. | Institution | 4,394,367 | 7.48% | US$ 17.93M | 2026-03-31 |
| Biotechnology Value Fund L.P. | Institution | 3,139,797 | 5.35% | US$ 12.81M | 2026-03-31 |
| Raymond James Financial, Inc. | Institution | 2,602,382 | 4.43% | US$ 10.62M | 2026-03-31 |
| BlackRock, Inc. | Institution | 2,364,866 | 4.03% | US$ 9.65M | 2026-03-31 |
| Biotechnology Value Fund II, L.P. | Institution | 2,218,382 | 3.78% | US$ 9.05M | 2026-03-31 |
| Flagship Pioneering Special Opportunities Fund II, L.P. | Institution | 1,851,801 | 3.15% | US$ 7.56M | 2026-01-13 |
| Euclidean Capital LLC | Institution | 1,573,761 | 2.68% | US$ 6.42M | 2026-03-31 |
| Vanguard Capital Management, LLC | Institution | 1,495,089 | 2.55% | US$ 6.10M | 2026-03-31 |
| Flagship Ventures Opportunities Fund I, L.P. | Institution | 1,491,441 | 2.54% | US$ 6.09M | 2026-01-13 |
| Geode Capital Management, LLC | Institution | 848,172 | 1.44% | US$ 3.46M | 2026-03-31 |
| Artal Group S.A. | Institution | 720,720 | 1.23% | US$ 2.94M | 2026-03-31 |
| State Street Global Advisors, Inc. | Institution | 679,787 | 1.16% | US$ 2.77M | 2026-03-31 |
| Columbia Management Investment Advisers, LLC | Institution | 448,976 | 0.76% | US$ 1.83M | 2026-03-31 |
| Renaissance Technologies LLC | Institution | 406,425 | 0.69% | US$ 1.66M | 2026-03-31 |
| Biotechnology Value Trading Fund OS LP | Institution | 402,467 | 0.69% | US$ 1.64M | 2026-03-31 |
| Candriam, Société En Commandite Par Actions | Institution | 313,765 | 0.53% | US$ 1.28M | 2026-03-31 |
| Jacobs Levy Equity Management Inc | Institution | 303,757 | 0.52% | US$ 1.24M | 2026-03-31 |
| Birchview Capital, LP | Institution | 281,730 | 0.48% | US$ 1.15M | 2026-03-31 |
| Northern Trust Global Investments | Institution | 275,875 | 0.47% | US$ 1.13M | 2026-03-31 |
| Vanguard Portfolio Management, LLC | Institution | 255,931 | 0.44% | US$ 1.04M | 2026-03-31 |
| Vanguard Fiduciary Trust Company | Institution | 240,586 | 0.41% | US$ 981.59k | 2026-03-31 |
| Citadel Advisors LLC | Institution | 200,013 | 0.34% | US$ 816.05k | 2026-03-31 |
| Woodline Partners LP | Institution | 180,333 | 0.31% | US$ 735.76k | 2026-03-31 |
| Sio Capital Management, LLC | Institution | 122,889 | 0.21% | US$ 501.39k | 2026-03-31 |
| PanAgora Asset Management, Inc. | Institution | 115,063 | 0.20% | US$ 469.46k | 2026-03-31 |
| Dafna Capital Management, LLC | Institution | 111,859 | 0.19% | US$ 456.38k | 2026-03-31 |
| SCS Capital Management LLC | Institution | 106,980 | 0.18% | US$ 436.48k | 2026-03-31 |
| Squarepoint OPS LLC | Institution | 103,743 | 0.18% | US$ 423.27k | 2026-03-31 |
| Charles Schwab Investment Management, Inc. | Institution | 103,525 | 0.18% | US$ 422.38k | 2026-03-31 |
| The B Group, Inc. | Institution | 102,000 | 0.17% | US$ 416.16k | 2026-03-31 |
| Morgan Stanley | Institution | 97,222 | 0.17% | US$ 396.67k | 2026-03-31 |
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